Plant tissues are divided into growth (embryonic) tissues, supporting, filling and photosynthetic tissues (ground tissues), conducting (vascular) tissues and covering (dermal) tissues.
Embryonic tissues: primary meristems; secondary meristems. Supporting tissues: collenchyma; sclerenchyma. Filling and photosynthetic tissues: photosynthetic parenchyma; storage parenchyma. Conducting tissues: xylem; phloem. Covering tissues: epidermis; periderm.
The growth tissues of the plants are meristems. Meristems are the tissues that produce plant growth, and are the origin of all other tissues. They are formed of undifferentiated cells with an intense cell division rate. Meristems are classified as primary meristems and secondary meristems.
Primary meristems are found at the apex of the stem, in the lateral buds of the stem, at the base and the tips of shoots and within the root cap. Primary meristems are responsible for the primary growth (lengthening) of the plant.
Secondary meristems make the plant grow in thickness (secondary growth) and are formed by tissues that thicken the stem: cambium and phellogen (cork cambium).
Lateral buds are portions of meristematic tissue located at the base of the shoots. Apical buds are portions of meristematic tissue situated at the tip of the stem and shoots.
Apical meristems are primary meristems found at the apex of the stem and in the tips of shoots and roots.
Apical meristems are responsible for the primary growth of plants.
Lateral, or secondary, meristems, consist of the cambium and the phellogen (also known as cork cambium). These are tissues in the stem, branches and roots that generate other tissues via mitosis. These tissues participate in the secondary growth of plant, that is, in the thickening of the stem, branches and roots.
Meristematic cells have very thin cell walls, small vacuoles, a well-centralized nucleus and they are constantly undergoing mitosis. Meristematic cells need a high mitotic rate because they are responsible for plant growth.
This type of tissue is most likely meristematic tissue. Meristematic tissues, when seen under a microscope, contain a large number of cells undergoing mitosis.
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The supportive tissues of plants are collenchyma and sclerenchyma.
Collenchyma is made of elongated, living cells that accumulate cellulose and pectin in some regions of the cell wall,giving them an unequal thickness and thus providing flexibility.
Sclerenchyma mostly consists of dead cells killed by lignin deposits (lignin is an impermeable biopolymer), which form elongated, rigid and impermeable fibers. Sclerenchyma is a plant tissue widely used in the textile industry.
The main photosynthetic tissue is photosynthetic parenchyma (also known as chlorenchyma, do not to be confused with collenchyma), which is often located between the superior and the inferior epidermis of the leaves.
Water, mineral salts and sugar are transported throughout the plant through conducting vessels formed by specialized tissues.
Xylem and Phloem are the vascular tissues of plants. Xylem is the plant tissue that form the vessels that transport water and mineral salts absorbed from the soil to plant cells. Phloem is the plant tissue that forms the vessels that transport dissolved sugar from the leaves (where it is produced via photosynthesis) to other plant cells.
The main cells of xylem are tracheids and vessel elements (these are only in found in angiosperms). Tracheids and vessel elements are dead cells that have lost their cytoplasm. Only their cell wall impregnated with lignin (an impermeable biopolymer) remains. Tracheids form tubes that connect to neighboring tubes through pores. Vessel elements do not contain pores but instead they connect to the following vessel element through perforations at their extremities.
The main cells that form phloem are sieve elements and companion cells. Sieve elements form the vessel walls. They are living enucleated cells positioned in series to form sieve tubes. Between successive vessel elements are communicating pores. Companion cells are located outside and alongside the sieve tubes and they help in the absorption of the material to be transported.
The vascular cambium is the secondary meristematic tissue that forms the vascular tissues (xylem and phloem) of the plant in roots and in the stem. Usually, the outer side of the vascular cambium produces a layer of phloem and the inner (more central) side of the tissue produces a layer of xylem.
Vascular bundles are segments of xylem attached to phloem that run longitudinally within the stem. In dicots, vascular bundles are organized side-by-side forming concentric rings. In monocots, vascular bundles are scattered and do not form rings.
The rings observed on a trunk cross section of dicot trees are made of conducting tissues: xylem and phloem.
For tree to grow, it is necessary new vessels within the stem to be formed, a task performed by the vascular cambium. The vascular cambium is more active during hot seasons (summer and spring), generating a lighter band made of large-diameter vessels. During winter and fall, the vascular cambium produces the opposite. As a result, small-diameter vessels and a darker band appears around the previous lighter band. Therefore two rings are made per year, one of which is lighter and the other of which is darker. By counting these pairs directly, the age of the tree can be estimated.
Leaf veins are made of vascular tissues. They are composed of xylem and phloem, which respectively conduct water and mineral nutrients (xylem) and sugar (phloem).
Covering tissues, or dermal tissues, in plants are the epidermis (which covers the leaves, the young stems and shoots) and the periderm (a tissue that replaces the epidermis in stems, shoots and roots). The periderm is made of phelloderm, phellogen and suber (cork).
The stem may be covered by epidermis (which contains stomata, cuticle and photosynthetic cells) as is the case in monocots or, alternatively, epidermis may be replaced by periderm (phelloderm, phellogen and cork) as is the case in dicots and gymnosperms.
Leaves are covered by epidermis.
Phellogen, also known as cork cambium, is the meristematic plant tissue responsible for the formation of periderm (the covering of the stem, shoots and roots). The inner side of the layer of phellogen forms the phelloderm and its outer side forms the cork (suber). The suber secretes suberin, an impermeable substance that enters the tissue.
Cork, the same material used as the cork of wine bottles, is extracted from the suber of a special oak tree called cork oak.
Root hairs are external elongated projections of the root epidermis. Their role is to increase the absorption of water by the root.
Bark is the mature periderm of the stem, branches and roots. It dies and breaks off when these structures grow, thus rupturing the peridermal suber formed of already dead cells.
The leaf cuticle is a thin waxy layer made of cutin and waxes, which is located on the outer surface of the leaf epidermis. Its function is to control cellular transpiration.
Roots have a central portion filled with a substance called medulla, which is made of vascular tissue (inner xylem and outer phloem). The medulla is surrounded by the medullary parenchyma and is enclosed by pericycle, a meristem that produces the secondary roots (ramifications). On the outside of the medulla lies the cortical portion, which is formed of endodermis (which surrounds the pericycle) and cortical parenchyma. The covering of the roots is epidermis (with root hairs), which is later replaced with suberized (corky) periderm.
The root cap is a protective structure located at the tip of a growing root. It protects the meristematic tissue of the root, forming a cap that surrounds its tip. This cover is necessary because during the growth of the root, the meristem would be injured by friction with the soil.
Secondary roots are branches of the primary (main) root. Secondary roots emerge from the pericycle, the inner tissue of the root. Shoots originate from the lateral buds of the stem. Therefore, the origin of secondary roots is endogenous and the origin of shoots is exogenous.
Swamp and marsh plants generally contain supporting roots that branch off from portions of the stem above the ground, thus helping the plant to establish itself in muddy and sandy soil. They may also contain respiratory roots (pneumatophores), structures that emerge from buried roots to absorb oxygen.
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